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Interleukin-6 is critical in the development of pulmonary vascular disease in Gcn2-deficient mice.

Title: Interleukin-6 is critical in the development of pulmonary vascular disease in Gcn2-deficient mice.
Authors: Schwiening, Max; Gao, Qingyue; Southwood, Mark; Crosby, Alexi; Moore, Stephen; Valer, Jose A; Veale, Niki; Dunmore, Benjamin J; Upton, Paul D; Thompson, AA Roger; Morrell, Nicholas W; Marciniak, Stefan J; Soon, Elaine
Publisher Information: National Academy of Sciences; C.I.M.R. Medicine; //doi.org/10.1073/pnas.2531623123
Publication Year: 2026
Collection: Apollo - University of Cambridge Repository
Subject Terms: GCN2
Description: Biallelic mutations in EIF2AK4, encoding Eukaryotic Translation Initiation Factor 2α kinase 4 or General Control Nonderepressible 2 (GCN2), cause pulmonary veno-occlusive disease (PVOD), a fatal form of pulmonary hypertension. The mechanisms linking GCN2 deficiency with pulmonary vascular pathology are poorly understood. To investigate this, we developed two mouse models: genetic ablation of Gcn2, to mirror GCN2-mutation positive PVOD, and a pharmacological model using mitomycin C, a drug which can cause PVOD as an idiosyncratic drug reaction. Both models were phenotyped, and lungs from wild-type and Gcn2-deficient mice were analyzed using single-cell RNA sequencing. We show that homozygous loss of Gcn2 is sufficient to induce mild pulmonary hypertension in mice. Single-cell transcriptomic profiling identified adventitial fibroblasts as the cell population exhibiting the most Gcn2-dependent transcriptional changes. Pathway analysis revealed upregulation of inflammatory signaling in Gcn2-/- adventitial fibroblasts. Consistent with this, we demonstrate a proinflammatory phenotype in Gcn2-/- mouse fibroblasts and in Gcn2-/- mice. Using a mitomycin C-induced murine model, genetic deletion of interleukin-6 (Il6) rescued the pulmonary vascular phenotype. Furthermore, chronic lipopolysaccharide exposure exaggerated pulmonary hypertension in Gcn2-/- mice, and Il6 ablation rescued both baseline and lipopolysaccharide-exacerbated disease. Pharmacological inhibition or genetic ablation of the Integrated Stress Response, which can be driven by GCN2-activation, phenocopies Gcn2 deficiency. Therefore, we establish a regulatory effect of an intact GCN2-Integrated Stress Response on IL-6 signaling. Together, we show that interleukin-6 is a critical mediator of both Gcn2 deficiency-associated and mitomycin C-triggered pulmonary vascular disease in mice and highlight IL-6-dependent pathways as potential therapeutic targets.
Document Type: article in journal/newspaper
File Description: application/pdf
Language: English
Relation: https://www.repository.cam.ac.uk/handle/1810/404205; https://doi.org/10.17863/CAM.130899
DOI: 10.17863/CAM.130899
Availability: https://www.repository.cam.ac.uk/handle/1810/404205; https://doi.org/10.17863/CAM.130899
Rights: Attribution 4.0 International ; https://creativecommons.org/licenses/by/4.0/
Accession Number: edsbas.3AB03E67
Database: BASE