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Germinal Center Hypoxia Potentiates Immunoglobulin Class Switch Recombination

Title: Germinal Center Hypoxia Potentiates Immunoglobulin Class Switch Recombination
Authors: Abbott, Robert K; Thayer, Molly; Labuda, Jasmine; Silva, Murillo; Philbrook, Phaethon; Cain, Derek W; Kojima, Hidefumi; Hatfield, Stephen; Sethumadhavan, Shalini; Ohta, Akio; Reinherz, Ellis L; Kelsoe, Garnett; Sitkovsky, Michail
Source: The Journal of Immunology ; volume 197, issue 10, page 4014-4020 ; ISSN 0022-1767 1550-6606
Publisher Information: Oxford University Press (OUP)
Publication Year: 2016
Description: Germinal centers (GCs) are anatomic sites where B cells undergo secondary diversification to produce high-affinity, class-switched Abs. We hypothesized that proliferating B cells in GCs create a hypoxic microenvironment that governs their further differentiation. Using molecular markers, we found GCs to be predominantly hypoxic. Compared to normoxia (21% O2), hypoxic culture conditions (1% O2) in vitro accelerated class switching and plasma cell formation and enhanced expression of GL-7 on B and CD4+ T cells. Reversal of GC hypoxia in vivo by breathing 60% O2 during immunization resulted in reduced frequencies of GC B cells, T follicular helper cells, and plasmacytes, as well as lower expression of ICOS on T follicular helper cells. Importantly, this reversal of GC hypoxia decreased Ag-specific serum IgG1 and reduced the frequency of IgG1+ B cells within the Ag-specific GC. Taken together, these observations reveal a critical role for hypoxia in GC B cell differentiation.
Document Type: article in journal/newspaper
Language: English
DOI: 10.4049/jimmunol.1601401
Availability: https://doi.org/10.4049/jimmunol.1601401; https://academic.oup.com/jimmunol/article-pdf/197/10/4014/61569708/1601401.pdf
Rights: https://academic.oup.com/pages/standard-publication-reuse-rights
Accession Number: edsbas.5A9ECEB4
Database: BASE