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Formin Is Associated with Left-Right Asymmetry in the Pond Snail and the Frog

Title: Formin Is Associated with Left-Right Asymmetry in the Pond Snail and the Frog
Authors: Davison, Angus; McDowell, Gary S.; Holden, Jennifer M.; Johnson, Harriet F.; Koutsovoulos, Georgios D.; Liu, M. Maureen; Hulpiau, Paco; van Roy, Frans; Wade, Christopher M.; Banerjee, Ruby; Yang, F.; Chiba, Satoshi; Davey, John W.; Jackson, Daniel John; Levin, Michael; Blaxter, Mark L.
Contributors: Davison, Angus; McDowell, Gary S.; Holden, Jennifer M.; Johnson, Harriet F.; Koutsovoulos, Georgios D.; Liu, M. Maureen; Hulpiau, Paco; van Roy, Frans; Wade, Christopher M.; Banerjee, Ruby; Yang, F.; Chiba, Satoshi; Davey, John W.; Jackson, Daniel John; Levin, Michael; Blaxter, Mark L.
Publisher Information: Cell Press
Publication Year: 2016
Collection: Georg-August-Universität Göttingen: GoeScholar
Description: While components of the pathway that establishes left-right asymmetry have been identified in diverse animals, from vertebrates to flies, it is striking that the genes involved in the first symmetry-breaking step remain wholly unknown in the most obviously chiral animals, the gastropod snails. Previously, research on snails was used to show that left-right signaling of Nodal, downstream of symmetry breaking, may be an ancestral feature of the Bilateria [1, 2]. Here, we report that a disabling mutation in one copy of a tandemly duplicated, diaphanous-related formin is perfectly associated with symmetry breaking in the pond snail. This is supported by the observation that an anti-formin drug treatment converts dextral snail embryos to a sinistral phenocopy, and in frogs, drug inhibition or overexpression by microinjection of formin has a chirality-randomizing effect in early (pre-cilia) embryos. Contrary to expectations based on existingmodels [3-5], we discovered asymmetric gene expression in 2- and 4-cell snail embryos, preceding morphological asymmetry. As the formin-actin filament has been shown to be part of an asymmetry-breaking switch in vitro [6, 7], together these results are consistent with the view that animals with diverse body plans may derive their asymmetries from the same intracellular chiral elements [8].
Document Type: article in journal/newspaper
Language: unknown
ISBN: 978-0-00-371401-2; 0-00-371401-2
Relation: 000371401200024
DOI: 10.1016/j.cub.2015.12.071
Availability: https://resolver.sub.uni-goettingen.de/purl?gro-2/41134; https://resolver.sub.uni-goettingen.de/purl?gs-1/14154; https://doi.org/10.1016/j.cub.2015.12.071
Rights: info:eu-repo/semantics/openAccess ; CC BY 4.0 ; https://creativecommons.org/licenses/by/4.0/
Accession Number: edsbas.5E35C918
Database: BASE