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Inhibition of ice recrystallization with designed twistless helical repeat proteins

Title: Inhibition of ice recrystallization with designed twistless helical repeat proteins
Authors: de Haas, Robbert J.; Pyles, Harley; Huddy, Evelyn B.; van Ossenbruggen, Jannick; Zheng, Chuanbao; van den Broek, Daniëlle; Giezen, Sanne N.; Carr, Ann; Bera, Asim K.; Kang, Alex; Brackenbrough, Evans; Joyce, Emily; Sankaran, Banumathi; Baker, David; Voets, Ilja K.; de Vries, Renko
Source: Proceedings of the National Academy of Sciences of the United States of America 122 (2025) 48 ; ISSN: 0027-8424
Publication Year: 2025
Collection: Wageningen UR (University & Research Centre): Digital Library
Subject Terms: ice binding proteins; ice recrystallization; protein design
Description: Given the repetitive structure of crystalline ice, it is unsurprising that highly active ice-binding proteins (IBPs), often with beta-roll structures, also have repeating motifs. Here, we introduce a de novo designed family of ice-binding twistless alpha-helical repeat (iTHR) proteins. Each iTHR protein comprises two planar layers of parallel alpha-helices connected by loops—a structural topology not seen in native IBPs. The ice-binding helices contain an ordered array of TXXXAXXXAXX motifs, precisely spaced to complement the pyramidal {201} and secondary prism {110} planes of the ice lattice, with a designed 98.2° residue turn angle that orients all threonines uniformly toward the ice surface. iTHR proteins show high solubility, thermostability, and produce varied ice crystal morphologies depending on their intended target facet. Crucially, iTHRs exhibit ice recrystallization inhibition (IRI) at critical concentrations comparable to those of many native globular IBPs. Extensive site-specific mutagenesis shows that ice-binding activity in iTHR proteins is robust, remaining largely unaffected by changes in chemical composition. Variation in the repeat number reveals a nonmonotonic relationship to IRI activity. X-ray crystal structures of two designs confirm the intended orientation of threonines, uniformly pointing toward the ice surface. The iTHR family provides a versatile platform to systematically investigate the complex structure–activity relationships underlying protein–ice interactions.
Document Type: article in journal/newspaper
File Description: application/pdf
Language: English
Relation: https://edepot.wur.nl/706106
DOI: 10.1073/pnas.2514871122
Availability: https://research.wur.nl/en/publications/inhibition-of-ice-recrystallization-with-designed-twistless-helic-2; https://doi.org/10.1073/pnas.2514871122; https://edepot.wur.nl/706106
Rights: https://creativecommons.org/licenses/by-nc-nd/4.0/ ; Wageningen University & Research
Accession Number: edsbas.71754703
Database: BASE