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HIV-1 infection regulates gene expression by altering alternative polyadenylation correlated with CPSF6 and CPSF5 redistribution

Title: HIV-1 infection regulates gene expression by altering alternative polyadenylation correlated with CPSF6 and CPSF5 redistribution
Authors: Charlotte Luchsinger; Annie Zhi Dai; Hari Yalamanchili; Aiswarya Balakrishnan; Kai-Lieh Huang; Cinzia Bertelli; Bin Cui; Ramon Lorenzo-Redondo; Eric J. Wagner; Felipe Diaz-Griffero
Source: mBio, Vol 17, Iss 1 (2026)
Publisher Information: American Society for Microbiology
Publication Year: 2026
Collection: Directory of Open Access Journals: DOAJ Articles
Subject Terms: HIV-1; CPSF6; CPSF5; capsid; alternative polyadenylation(APA); gene expression; Microbiology; QR1-502
Description: HIV-1 viral core transport to the nucleus, an early infection event, triggers the redistribution of cleavage and polyadenylation specificity factors (CPSF) 5 and CPSF6 to nuclear speckles, forming puncta-like structures. CPSF5 and CPSF6 regulate alternative polyadenylation (APA), which governs approximately 70% of gene expression. APA alters the lengths of mRNA 3’-untranslated regions (3′-UTRs), which contain regulatory signals influencing RNA stability, localization, and function. We investigated whether HIV-1 infection–induced changes in CPSF5 and CPSF6 subcellular localization are accompanied by APA changes. Using two independent methodologies to assess APA in human cell lines and primary CD4+ T cells, we found that HIV-1 infection regulates APA, shaped by the interaction of CPSF6 with the viral capsid, recapitulating the APA phenotype observed in CPSF6 knockout cells. Our study demonstrates that HIV-1 infection leverages the interaction between the viral capsid and CPSF6 to co-opt cellular processes, alter gene expression, and potentially contribute to viral pathogenesis.IMPORTANCEThe interaction between HIV-1 and the cellular protein CPSF6 has been known for over 15 years; however, depletion of CPSF6 does not impair productive infection. An alternative possibility is that the virus exploits this protein to modulate cellular processes. This study demonstrates that HIV-1 infection alters the cellular function of CPSF6, an essential regulator of alternative polyadenylation—a mechanism that controls 70% of gene expression. Here, we show that HIV-1 regulates gene expression by disrupting the alternative polyadenylation function of CPSF6 through direct interaction. Overall, this reveals a novel strategy employed by the virus to modulate cellular gene expression.
Document Type: article in journal/newspaper
Language: English
Relation: https://doaj.org/toc/2150-7511; https://doaj.org/article/2d2d137ae6464837bdd3984b6a5b2951
DOI: 10.1128/mbio.02865-25
Availability: https://doi.org/10.1128/mbio.02865-25; https://doaj.org/article/2d2d137ae6464837bdd3984b6a5b2951
Accession Number: edsbas.83D733D
Database: BASE