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A RAD18-UBC13-PALB2-RNF168 axis mediates replication fork recovery in BRCA1-deficient cancer cells

Title: A RAD18-UBC13-PALB2-RNF168 axis mediates replication fork recovery in BRCA1-deficient cancer cells
Authors: Cybulla, Emily; Wallace, Sierra; Meroni, Alice; Jackson, Jessica; Agashe, Sumedha; Tennakoon, Mithila; Limbu, Mangsi; Quinet, Annabel; Lomonosova, Elena; Noia, Hollie; Tirman, Stephanie; Wood, Matthew; Lemacon, Delphine; Fuh, Katherine; Zou, Lee; Vindigni, Alessandro
Source: Cybulla, Emily; Wallace, Sierra; Meroni, Alice; Jackson, Jessica; Agashe, Sumedha; Tennakoon, Mithila; Limbu, Mangsi; Quinet, Annabel; Lomonosova, Elena; Noia, Hollie; Tirman, Stephanie; Wood, Matthew; Lemacon, Delphine; Fuh, Katherine; Zou, Lee; Vindigni, Alessandro (2024). A RAD18-UBC13-PALB2-RNF168 axis mediates replication fork recovery in BRCA1-deficient cancer cells. Nucleic Acids Research, 52(15):8861-8879.
Publisher Information: Oxford University Press
Publication Year: 2024
Collection: University of Zurich (UZH): ZORA (Zurich Open Repository and Archive
Subject Terms: Clinic for Gynecology; 610 Medicine & health
Description: BRCA1/2 proteins function in genome stability by promoting repair of double-stranded DNA breaks through homologous recombination and by protecting stalled replication forks from nucleolytic degradation. In BRCA1/2-deficient cancer cells, extensively degraded replication forks can be rescued through distinct fork recovery mechanisms that also promote cell survival. Here, we identified a novel pathway mediated by the E3 ubiquitin ligase RAD18, the E2-conjugating enzyme UBC13, the recombination factor PALB2, the E3 ubiquitin ligase RNF168 and PCNA ubiquitination that promotes fork recovery in BRCA1- but not BRCA2-deficient cells. We show that this pathway does not promote fork recovery by preventing replication fork reversal and degradation in BRCA1-deficient cells. We propose a mechanism whereby the RAD18-UBC13-PALB2-RNF168 axis facilitates resumption of DNA synthesis by promoting re-annealing of the complementary single-stranded template strands of the extensively degraded forks, thereby allowing re-establishment of a functional replication fork. We also provide preliminary evidence for the potential clinical relevance of this novel fork recovery pathway in BRCA1-mutated cancers, as RAD18 is over-expressed in BRCA1-deficient cancers, and RAD18 loss compromises cell viability in BRCA1-deficient cancer cells.
Document Type: article in journal/newspaper
File Description: application/pdf
Language: English
ISSN: 0305-1048
Relation: https://www.zora.uzh.ch/id/eprint/270508/1/gkae563.pdf; info:pmid/38943334; urn:issn:0305-1048
DOI: 10.1093/nar/gkae563
Availability: https://www.zora.uzh.ch/id/eprint/270508/; https://www.zora.uzh.ch/id/eprint/270508/1/gkae563.pdf; https://doi.org/10.1093/nar/gkae563
Rights: info:eu-repo/semantics/openAccess ; Creative Commons: Attribution 4.0 International (CC BY 4.0) ; http://creativecommons.org/licenses/by/4.0/
Accession Number: edsbas.C67D5073
Database: BASE