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Human T-cell leukemia virus type 1 Tax and cell progression. Role of cyclin D-cdk and p110 Rb

Title: Human T-cell leukemia virus type 1 Tax and cell progression. Role of cyclin D-cdk and p110 Rb
Authors: NEUVEUT C; LOW KG; MALDARELLI F; SCHMITT I; GRASSMANN R; JEANG KT; MAJONE, FRANCA
Contributors: Neuveut, C; Low, Kg; Maldarelli, F; Schmitt, I; Majone, Franca; Grassmann, R; Jeang, Kt
Publisher Information: American Society for Microbiology / DC:1752 N Street Northwest:Washington, DC 20036:(202)737-3600, EMAIL: jadelman@asmusa.org, INTERNET: http://www.asmusa.org/asm.htm, Fax: (202)942-9342
Publication Year: 1998
Collection: Padua Research Archive (IRIS - Università degli Studi di Padova)
Subject Terms: RETINOBLASTOMA GENE-PRODUCT; DEPENDENT KINASE INHIBITORS; LONG TERMINAL REPEATS; HTLV-1 TAX
Description: Human T-cell leukemia virus type 1 is etiologically linked to the development of adult T-cell leukemia and various human neuropathies. The Tax protein of human T-cell leukemia virus type I has been implicated in cellular transformation. Like other oncoproteins, such as Myc, Jun, and Fos, Tax is a transcriptional activator. How it mechanistically dysregulates the cell cycle is unclear. Previously, it was suggested that Tax affects cell-phase transition by forming a direct protein-protein complex with p16(INK4a), thereby inactivating an inhibitor of G1-to-S-phase progression. Here we show that, in T cells deleted for p16(INK4a), Tax can compel an egress of cells from G0/G1 into S despite the absence of serum. We also show that in undifferentiated myocytes, expression of Tax represses cellular differentiation. In both settings, Tax expression was found to increase cyclin D-cdk activity and to enhance pRb phosphorylation. In T cells, a Tax-associated increase in steady-state E2F2 protein was also documented. In searching for a molecular explanation for these observations, we found that Tax forms a protein-protein complex with cyclin D3, whereas a point-mutated and transcriptionally inert Tax mutant failed to form such a complex. Interestingly, expression of wild-type Tax protein in cells was also correlated with the induction of a novel hyperphosphorylated cyclin D3 protein. Taken together, these findings suggest that Tax might directly influence cyclin D-cdk activity and function, perhaps by a route independent of cdk inhibitors such as p16(INK4a).
Document Type: article in journal/newspaper
File Description: STAMPA
Language: English
Relation: info:eu-repo/semantics/altIdentifier/pmid/9584203; info:eu-repo/semantics/altIdentifier/wos/WOS:000073628800053; volume:18; firstpage:3620; lastpage:3632; numberofpages:13; journal:MOLECULAR AND CELLULAR BIOLOGY; https://hdl.handle.net/11577/110128
Availability: https://hdl.handle.net/11577/110128
Rights: info:eu-repo/semantics/openAccess ; license:Accesso gratuito
Accession Number: edsbas.E7B6CBA8
Database: BASE