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Complex excitability and “flipping' of granule cells: An experimental and computational study

Title: Complex excitability and “flipping' of granule cells: An experimental and computational study
Authors: Danielewicz, Joanna; Girier, Guillaume; Chizhov, Anton; Desroches, Mathieu; Encinas, Juan-Manuel; Rodrigues, Serafim
Contributors: Kuznetsov, Alexey; Ministerio de Ciencia, Innovación y Universidades; Eusko Jaurlaritza
Source: PLOS One ; volume 21, issue 2, page e0339418 ; ISSN 1932-6203
Publisher Information: Public Library of Science (PLoS)
Publication Year: 2026
Collection: PLOS Publications (via CrossRef)
Description: In response to prolonged depolarizing current steps, different classes of neurons display specific firing characteristics (i.e., excitability class), such as a regular train of action potentials with more or less adaptation, delayed responses, or bursting. In general, one or more specific ionic transmembrane currents underlie the different firing patterns. Here, we sought to investigate the influence of artificial sodium-like (Na channels) and slow potassium-like (KM channels) voltage-gated channels conductances on firing patterns and transition to depolarization block (DB) in Dentate Gyrus granule cells with dynamic clamp - a computer-controlled real-time closed-loop electrophysiological technique, which allows to couple mathematical models simulated in a computer with biological cells. Our findings indicate that the mimicked extra Na/KM channels significantly affect the firing rate of low-frequency cells, but not that of high-frequency cells. Moreover, we have observed that 44 percent of recorded cells exhibited what we have called a “flipping” behavior. This means that these cells were able to overcome the DB and generate trains of action potentials at higher current injection steps. We have developed a mathematical model of “flipping" cells to explain this phenomenon. Based on our computational model, we conclude that the appearance of “flipping" is linked to the number of states for the sodium channel of the model.
Document Type: article in journal/newspaper
Language: English
DOI: 10.1371/journal.pone.0339418
Availability: https://doi.org/10.1371/journal.pone.0339418; https://dx.plos.org/10.1371/journal.pone.0339418
Rights: http://creativecommons.org/licenses/by/4.0/
Accession Number: edsbas.EEED417F
Database: BASE