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Gene discovery and polygenic prediction from a genome-wide association study of educational attainment in 1.1 million individuals

Title:	Gene discovery and polygenic prediction from a genome-wide association study of educational attainment in 1.1 million individuals
Authors:	Lee, James J.^Aff1; Wedow, Robbee^{Aff2, Aff3, Aff4}; Okbay, Aysu^{Aff5, Aff6, cor3}; Kong, Edward^Aff7; Maghzian, Omeed^Aff7; Zacher, Meghan^Aff8; Nguyen-Viet, Tuan Anh^Aff9; Bowers, Peter^Aff7; Sidorenko, Julia^{Aff10, Aff11}; Karlsson Linnér, Richard^{Aff5, Aff6, Aff12}; Fontana, Mark Alan^{Aff9, Aff13}; Kundu, Tushar^Aff9; Lee, Chanwook^Aff7; Li, Hui^Aff7; Li, Ruoxi^Aff9; Royer, Rebecca^Aff9; Timshel, Pascal N.^{Aff14, Aff15}; Walters, Raymond K.^{Aff16, Aff17}; Willoughby, Emily A.^Aff1; Yengo, Loïc^Aff10; Alver, Maris^Aff11; Bao, Yanchun^Aff20; Clark, David W.^Aff21; Day, Felix R.^Aff22; Furlotte, Nicholas A.^Aff23; Joshi, Peter K.^{Aff21, Aff24}; Kemper, Kathryn E.^Aff10; Kleinman, Aaron^Aff23; Langenberg, Claudia^Aff22; Mägi, Reedik^Aff11; Trampush, Joey W.^{Aff25, Aff26}; Verma, Shefali Setia^Aff27; Wu, Yang^Aff10; Lam, Max^{Aff28, Aff29}; Zhao, Jing Hua^Aff22; Zheng, Zhili^{Aff10, Aff30}; Boardman, Jason D.^{Aff2, Aff3, Aff4}; Campbell, Harry^Aff21; Freese, Jeremy^Aff31; Harris, Kathleen Mullan^{Aff32, Aff33}; Hayward, Caroline^Aff34; Herd, Pamela^{Aff20, Aff35}; Kumari, Meena^Aff20; Lencz, Todd^{Aff36, Aff37, Aff38}; Luan, Jian’an^Aff22; Malhotra, Anil K.^{Aff36, Aff37, Aff38}; Metspalu, Andres^{Aff11, Aff39}; Milani, Lili^Aff11; Ong, Ken K.^Aff22; Perry, John R. B.^Aff22; Porteous, David J.^Aff40; Ritchie, Marylyn D.^Aff27; Smart, Melissa C.^Aff21; Smith, Blair H.^{Aff41, Aff42}; Tung, Joyce Y.^Aff23; Wareham, Nicholas J.^Aff22; Wilson, James F.^{Aff21, Aff34}; Beauchamp, Jonathan P.^Aff43; Conley, Dalton C.^Aff44; Esko, Tõnu^Aff11; Lehrer, Steven F.^{Aff45, Aff46, Aff47}; Magnusson, Patrik K. E.^Aff48; Oskarsson, Sven^Aff49; Pers, Tune H.^{Aff14, Aff15}; Robinson, Matthew R.^{Aff10, Aff50}; Thom, Kevin^Aff51; Watson, Chelsea^Aff9; Chabris, Christopher F.^Aff52; Meyer, Michelle N.^Aff53; Laibson, David I.^Aff7; Yang, Jian^{Aff10, Aff54}; Johannesson, Magnus^Aff55; Koellinger, Philipp D.^{Aff5, Aff6, Aff12}; Turley, Patrick^{Aff16, Aff17}; Visscher, Peter M.^{Aff10, Aff54, cor75}; Benjamin, Daniel J.^{Aff9, Aff47, Aff56, cor76}; Cesarini, David^{Aff47, Aff51, Aff57}; Agee, Michelle^Aff23; Alipanahi, Babak^Aff23; Auton, Adam^Aff23; Bell, Robert K.^Aff23; Bryc, Katarzyna^Aff23; Elson, Sarah L.^Aff23; Fontanillas, Pierre^Aff23; Hinds, David A.^Aff23; McCreight., Jennifer C.^Aff23; Huber, Karen E.^Aff23; Litterman, Nadia K.^Aff23; McIntyre, Matthew H.^Aff23; Mountain, Joanna L.^Aff23; Noblin, Elizabeth S.^Aff23; Northover, Carrie A. M.^Aff23; Pitts, Steven J.^Aff23; Sathirapongsasuti, J. Fah^Aff23; Sazonova, Olga V.^Aff23; Shelton, Janie F.^Aff23; Shringarpure, Suyash^Aff23; Tian, Chao^Aff23; Vacic, Vladimir^Aff23; Wilson, Catherine H.^Aff23; Okbay, Aysu^{Aff5, Aff6}; Beauchamp, Jonathan P.^Aff43; Lee, James J.^Aff1; Rietveld, Cornelius A.^{Aff12, Aff60, Aff61}; Chen, Guo-Bo^Aff54; Emilsson, Valur^{Aff62, Aff63}; Meddens, S. Fleur W.^{Aff5, Aff12, Aff64}; Pickrell, Joseph K.^Aff65; Timshel, Pascal^{Aff14, Aff15}; Vlaming, Ronald de^{Aff12, Aff60, Aff61}; Abdellaoui, Abdel^Aff66; Ahluwalia, Tarunveer S.^{Aff14, Aff67, Aff68}; Bacelis, Jonas^Aff69; Baumbach, Clemens^{Aff70, Aff71}; Bjornsdottir, Gyda^Aff72; Brandsma, Johannes H.^Aff73; Concas, Maria Pina^Aff74; Derringer, Jaime^Aff75; Galesloot, Tessel E.^Aff76; Girotto, Giorgia^Aff77; Gupta, Richa^Aff78; Hall, Leanne M.^{Aff79, Aff80}; Harris, Sarah E.^{Aff40, Aff80}; Hofer, Edith^{Aff81, Aff82}; Horikoshi, Momoko^{Aff83, Aff84}; Huffman, Jennifer E.^Aff34; Kaasik, Kadri^Aff85; Kalafati, Ioanna P.^Aff86; Karlsson, Robert^Aff48; Kong, Augustine^Aff72; Lahti, Jari^{Aff85, Aff87}; van der Lee, Sven J.^Aff61; Leeuw, Christiaan de^{Aff5, Aff88}; Lind, Penelope A.^Aff89; Lindgren, Karl-Oskar^Aff49; Liu, Tian^Aff90; Mangino, Massimo^{Aff91, Aff92}; Marten, Jonathan^Aff34; Mihailov, Evelin^Aff11; Miller, Michael B.^Aff1; van der Most, Peter J.^Aff93; Oldmeadow, Christopher^{Aff94, Aff95}; Payton, Antony^{Aff96, Aff97}; Pervjakova, Natalia^{Aff11, Aff98}; Peyrot, Wouter J.^Aff99; Qian, Yong^Aff100; Raitakari, Olli^Aff101; Rueedi, Rico^{Aff102, Aff103}; Salvi, Erika^Aff104; Schmidt, Börge^Aff105; Schraut, Katharina E.^Aff21; Shi, Jianxin^Aff106; Smith, Albert V.^{Aff62, Aff107}; Poot, Raymond A.^Aff73; St Pourcain, Beate^{Aff108, Aff109}; Teumer, Alexander^Aff110; Thorleifsson, Gudmar^Aff72; Verweij, Niek^Aff111; Vuckovic, Dragana^Aff77; Wellmann, Juergen^Aff112; Westra, Harm-Jan^{Aff113, Aff114, Aff115}; Yang, Jingyun^{Aff116, Aff117}; Zhao, Wei^Aff118; Zhu, Zhihong^Aff54; Alizadeh, Behrooz Z.^{Aff93, Aff119}; Amin, Najaf^Aff61; Bakshi, Andrew^Aff54; Baumeister, Sebastian E.^{Aff110, Aff120}; Biino, Ginevra^Aff121; Bønnelykke, Klaus^Aff67; Boyle, Patricia A.^{Aff116, Aff122}; Cappuccio, Francesco P.^Aff123; Davies, Gail^{Aff80, Aff124}; De Neve, Jan-Emmanuel^Aff125; Deloukas, Panos^{Aff126, Aff127}; Demuth, Ilja^{Aff128, Aff129}; Ding, Jun^Aff100; Eibich, Peter^{Aff130, Aff131}; Eisele, Lewin^Aff105; Eklund, Niina^Aff98; Evans, David M.^{Aff108, Aff132}; Faul, Jessica D.^Aff133; Feitosa, Mary F.^Aff134; Forstner, Andreas J.^{Aff135, Aff136}; Gandin, Ilaria^Aff77; Gunnarsson, Bjarni^Aff72; Halldórsson, Bjarni V.^{Aff72, Aff137}; Harris, Tamara B.^Aff138; Heath, Andrew C.^Aff139; Hocking, Lynne J.^Aff140; Holliday, Elizabeth G.^{Aff94, Aff95}; Homuth, Georg^Aff141; Horan, Michael A.^Aff142; Hottenga, Jouke-Jan^Aff66; de Jager, Philip L.^{Aff115, Aff143, Aff144}; Joshi, Peter K.^{Aff21, Aff24}; Jugessur, Astanand^Aff145; Kaakinen, Marika A.^Aff146; Kähönen, Mika^{Aff147, Aff148}; Kanoni, Stavroula^Aff126; Keltigangas-Järvinen, Liisa^Aff85; Kiemeney, Lambertus A. L. M.^Aff76; Kolcic, Ivana^Aff149; Koskinen, Seppo^Aff98; Kraja, Aldi T.^Aff134; Kroh, Martin^Aff130; Kutalik, Zoltan^{Aff102, Aff103, Aff150}; Latvala, Antti^Aff78; Launer, Lenore J.^Aff151; Lebreton, Maël P.^{Aff64, Aff152}; Levinson, Douglas F.^Aff153; Lichtenstein, Paul^Aff48; Lichtner, Peter^Aff154; Liewald, David C. M.^{Aff80, Aff124}; Loukola, LifeLines Cohort Study, Anu^Aff78; Madden, Pamela A.^Aff139; Mäki-Opas, Tomi^Aff98; Marioni, Riccardo E.^{Aff43, Aff80, Aff155}; Marques-Vidal, Pedro^Aff156; Meddens, Gerardus A.^Aff157; McMahon, George^Aff108; Meisinger, Christa^Aff71; Meitinger, Thomas^Aff154; Milaneschi, Yusplitri^Aff99; Milani, Lili^Aff11; Montgomery, Grant W.^Aff158; Myhre, Ronny^Aff145; Nelson, Christopher P.^{Aff79, Aff159}; Nyholt, Dale R.^{Aff158, Aff160}; Ollier, William E. R.^Aff96; Palotie, Aarno^{Aff16, Aff17, Aff115, Aff161, Aff162, Aff163}; Paternoster, Lavinia^Aff108; Pedersen, Nancy L.^Aff48; Petrovic, Katja E.^Aff81; Räikkönen, Katri^{Aff85, Aff87}; Ring, Susan M.^Aff108; Robino, Antonietta^Aff164; Rostapshova, Olga^{Aff7, Aff165}; Rudan, Igor^Aff21; Rustichini, Aldo^Aff166; Salomaa, Veikko^Aff98; Sanders, Alan R.^{Aff167, Aff168}; Sarin, Antti-Pekka^{Aff162, Aff169}; Schmidt, Helena^{Aff81, Aff170}; Scott, Rodney J.^{Aff95, Aff171}; Smith, Blair H.^{Aff41, Aff42}; Smith, Jennifer A.^Aff93; Staessen, Jan A.^{Aff172, Aff173}; Steinhagen-Thiessen, Elisabeth^Aff128; Strauch, Konstantin^{Aff174, Aff175}; Terracciano, Antonio^Aff176; Tobin, Martin D.^Aff177; Ulivi, Sheila^Aff164; Vaccargiu, Simona^Aff74; Quaye, Lydia^Aff91; van Rooij, Frank J. A.^{Aff61, Aff178}; Venturini, Cristina^{Aff91, Aff92}; Vinkhuyzen, Anna A. E.^Aff54; Völker, Uwe^Aff141; Völzke, Henry^Aff110; Vonk, Judith M.^Aff93; Vozzi, Diego^Aff164; Waage, Johannes^{Aff67, Aff68}; Ware, Erin B.^{Aff118, Aff179}; Willemsen, Gonneke^Aff66; Attia, John R.^{Aff94, Aff95}; Bennett, David A.^{Aff116, Aff117}; Berger, Klaus^Aff111; Bertram, Lars^{Aff180, Aff181}; Bisgaard, Hans^Aff67; Boomsma, Dorret I.^Aff66; Borecki, Ingrid B.^Aff134; Bültmann, Ute^Aff182; Cucca, Francesco^Aff183; Cusi, Daniele^{Aff104, Aff184}; Deary, Ian J.^{Aff80, Aff124}; Dedoussis, George V.^Aff86; van Duijn, Cornelia M.^Aff61; Eriksson, Johan G.^{Aff87, Aff185}; Franke, Barbara^Aff186; Franke, Lude^Aff187; Gasparini, Paolo^{Aff77, Aff164, Aff188}; Gejman, Pablo V.^{Aff167, Aff168}; Gieger, Christian^Aff70; Grabe, Hans-Jörgen^{Aff189, Aff190}; Gratten, Jacob^Aff54; Groenen, Patrick J. F.^Aff191; Gudnason, Vilmundur^{Aff62, Aff107}; van der Harst, Pim^{Aff111, Aff187, Aff192}; Hayward, Caroline^Aff34; Hoffmann, Wolfgang^Aff110; Hyppönen, Elina^{Aff193, Aff194, Aff195}; Iacono, William G.^Aff1; Jacobsson, Bo^{Aff69, Aff145}; Järvelin, Marjo-Riitta^{Aff196, Aff197, Aff198, Aff199}; Jöckel, Karl-Heinz^Aff105; Kaprio, Jaakko^{Aff78, Aff98, Aff162}; Kardia, Sharon L. R.^Aff118; Lehtimäki, Terho^{Aff200, Aff201}; Magnusson, Patrik K. E.^Aff48; Martin, Nicholas G.^Aff202; McGue, Matt^Aff1; Pendleton, Neil^{Aff203, Aff204}; Penninx, Brenda W. J. H.^Aff99; Perola, Markus^{Aff11, Aff98}; Pirastu, Nicola^Aff77; Pirastu, Mario^Aff74; Polasek, Ozren^{Aff21, Aff205}; Posthuma, Danielle^{Aff5, Aff206}; Power, Christine^Aff194; Province, Michael A.^Aff134; Samani, Nilesh J.^{Aff79, Aff159}; Schlessinger, David^Aff100; Schmidt, Reinhold^Aff81; Sørensen, Thorkild I. A.^{Aff14, Aff108, Aff207}; Spector, Tim D.^Aff91; Stefansson, Kari^{Aff72, Aff107}; Thorsteinsdottir, Unnur^{Aff72, Aff107}; Thurik, A. Roy^{Aff12, Aff60, Aff208, Aff209}; Timpson, Nicholas J.^Aff108; Tiemeier, Henning^{Aff61, Aff210, Aff211}; Uitterlinden, André G.^{Aff61, Aff212}; Vitart, Veronique^Aff34; Vollenweider, Peter^Aff156; Weir, David R.^Aff133; Wilson, James F.^{Aff21, Aff34}; Wright, Alan F.^Aff34; Krueger, Robert F.^Aff1; Smith, George Davey^Aff108; Hofman, Albert^Aff61; Medland, Sarah E.^Aff89; Yang, Jian^{Aff10, Aff54}; Johannesson, Magnus^Aff55; Visscher, Peter M.^{Aff10, Aff54}; Benjamin, Daniel J.^{Aff9, Aff47, Aff56}
Source:	Nature Genetics. 50(8):1112-1121
Database:	Springer Nature Journals